On the Cover: Anthers develop from undifferentiated meristematic cells into an organized set of tissues that produce pollen in plants. The cover image shows a high-pressure frozen freeze-substituted pre-meiotic barley anther including middle layer (bright green), tapetum (yellow-green), nuclei (blue) and mitochondria (red). Original article by Bélanger et al. provides novel developmental insights and evolutionary significance of phasiRNA biogenesis in maize, barley and rice. Image credit: Sébastien Bélanger & Kirk J Czymmek.