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On the Cover: Nonselective cation channels (NSCs) have been characterized in root plasma membranes from Arabidopsis (top right, Demidchik and Tester, pp. 379-387) and biotrophic interfaces in other plants. One class of these channels in Arabidopsis roots is gated by cyclic nucleotides (Maathuis and Sanders, Vol. 127, pp. 1617-1625). The ribbon diagram in the bottom left is for a cyclic nucleotide-gated channel (AtCNGC2) pore domain with the four subunits oriented to form the membrane-traversing pore (from Richard W. Mercier and Gerald A. Berkowitz; see also Leng et al., pp. 400-410). In biotrophic interfaces, nonselective channels are proposed to function in the release of nutrient cations (Zhang et al., pp. 388-399; Roberts and Tyerman, pp. 370-378). Design by Stephen D. Tyerman with confocal image of a root of Arabidopsis (propidium iodide stained) provided by Andrew Baker (Plant Sciences, Cambridge University, UK).