Table I.

Comparison of photosynthetic characteristics in three amphibious species of Eleocharis

Characteristics	E. vivipara		E. baldwinii		E. retroflexa
Characteristics	Terrestrial	Submerged	Terrestrial	Submerged	Terrestrial	Submerged
Culm anatomy	Kranz	Non-Kranz	Kranz	Non-Kranz-like	Kranz	Kranz-like
Organelles in bundle sheath cells	Prominent	Few	Prominent	Present	Prominent	Prominent^1-a
¹⁴C pulse-¹²C chase pattern	C₄	C₃	C₄	Intermediate	C₄	C₄
C₄enzymes	High^1-b	Low	High^1-b	Intermediate	High^1-b	High
δ¹³C values^1-c	C₄	–	C₄	–	C₄	–

↵F1-a There are fewer organelles than in the terrestrial form.
↵F1-b There is a gradient in the activities and amounts of C₄ enzymes among the terrestrial forms of the three species. See text for details.
↵F1-c Range of δ¹³C values is shown only for the terrestrial forms. The δ¹³C values of the terrestrial form of E. vivipara are slightly more negative than those of other terrestrial forms.